Extinction Episodes of the Past
It was not until the Cambrian period, beginning about 600 million years ago, that a great proliferation of macroscopic species occurred on Earth and produced a fossil record that allows us to track the rise and fall of biodiversity. Since the Cambrian period, biodiversity has generally risen, but there have been some notable exceptions. Biodiversity collapsed dramatically during at least five periods because of mass extinctions around the globe. The five major mass extinctions receive most of the attention, but they are only one end of a spectrum of extinction events. Collectively, more species went extinct during smaller events that were less dramatic but more frequent. The best known of the five major extinction events, the one that saw the demise of the dinosaurs, is the Cretaceous-Tertiary extinction.
Starting about 280 million years ago, reptiles were the dominant large animals in terrestrial environments. In popular language this was the era “when dinosaurs ruled Earth,” with a wide variety of reptile species occupying many ecological niches. However, no group or species can maintain its dominance indefinitely, and when, after over 200 million years, the age of dinosaurs came to a dramatic end about 65 million years ago, mammals began to flourish, evolving from relatively few types of small terrestrial animals into the myriad of diverse species, including bats and whales, that we know today. Paleontologists label this point in Earth’s history as the end of the Cretaceous period and the beginning of the Tertiary period, often abbreviated as the K-T boundary. This time was also marked by changes in many other types of organisms. Overall, about 38 percent of the families of marine animals were lost, with percentages much higher in some groups. Ammonoid mollusks went from being very diverse and abundant to being extinct. An extremely abundant set of planktonic marine animals called foraminifera largely disappeared, although they rebounded later. Among plants, the K-T boundary saw a sharp but brief rise in the abundance of primitive vascular plants such as ferns, club mosses, horsetails, and conifers and other gymnosperms. The number of flowering plants (angiosperms) was reduced at this time, but they then began to increase dramatically.
What caused these changes? For many years scientists assumed that a cooling of the climate was responsible, with dinosaurs being particularly vulnerable because, like modern reptiles, they were ectothermic (dependent on environmental heat, or cold-blooded). It is now widely believed that at least some species of dinosaurs had a metabolic rate high enough for them to be endotherms (animals that maintain a relatively consistent body temperature by generating heat internally). Nevertheless, climatic explanations for the K-T extinction are not really challenged by the idea that dinosaurs may have been endothermic, because even endotherms can be affected by a significant change in the climate.
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